Peer-reviewed papers


Murray, C., Markager, S., Stedmon, C.A., Juul-Pedersen, T., Sejr, M.K. and Bruhn, A. 2015. The influence of glacial melt water on bio-optical properties in two contrasting Greenland fjords. Estuarine, Coastal and Shelf Science. Doi:10.1016/j.ecss.2015.05.041.


Attard, K.M., Glud, R.N., McGinnis, D.F. and Rysgaard, S. 2014. Seasonal rates of benthic primary production in a Greenland fjord measured by aquatic eddy correlation. Limnology Oceanography 59(5):1555-1569.

Riisgaard, K. et al 2014. Trophic role and top-down control of a subarctic protozooplankton community. Marine Ecology Progress Series 500:67-82.

Sejr, M.K., Krause-Jensen, D., Dalsgaard, T., Halpern, S., Duarte, C.M., Middelboe, M., Glud, R.N., Rysgaard, S. and Bendtsen, J. 2014. Seasonal dynamics of autotrophic and heterotrophic plankton metabolism and pCO2 in a subarctic Greenland fjord. Limnology and Oceanography 59(5):1764-1778. doi:10.4319/lo.2014.59.5.1764.


Abermann, J., van As, D., Petersen, D. and Nauta, M. 2014. A new glacier monitoring site in West Greenland. Poster presented at the AGU meeting, San Francisco, USA. December 2014. 


Blicher, M.E., Høgslund, S. and Sejr, M.K. 2013. Population structure of Mytilus edulisin the intertidal zone in a sub-Arctic fjord, SW Greenland. Marine Ecology Progress Series 487:89-100.

Dziallas, C., Grossart, H.P., Tang, K.W. and Nielsen, T.G. 2013. Distinct Communities of Free-Living and Copepod-Associated Microorganisms along a Salinity Gradient in Godthåbsfjord, West Greenland. Arctic, Antarctic and Alpine Research 45(4):471-480.

Josefson, A.B., Mokievsky, V., Bergmann, M., Blicher, M.E., Bluhm, B., Cochrane, S., Denisenko, N.V., Hasemann, C., Jørgensen, L. L., Klages, M., Schewe, I., Sejr, M.K., Soltwedel, T., Weslawski, J.M. and Wlodarska-Kowalczuk, M. 2013. Marine Invertebrates p. 277-309. In Meltofte, H. (ed.) 2013. Arctic Biodiversity Assessment. Status and trends in Arctic biodiversity. Conservation of Arctic Flora and Fauna, Akureyri, Iceland. 674 pp. ISBN: 978-9935-431-22-6.

Kaartokallio, H., Søgaard, D., Norman, L., Rysgaard, S, Tison, J-L., Delille, B. and Thomas, D.N. 2013. Short- term variability in bacterial abundance, cell properties, leucine and thymidine incorporation in sub-Arctic sea ice. Aquatic Microbial Ecology 71:57-73.

Krawczyk, D.W., Witkowski, A., Lloyd, J., Moros, M., Harff, J. and Kuijpers, A. 2013. Late-Holocene diatom derived seasonal variability in hydrological conditions off Disko Bay, West Greenland. Quaternary Science Reviews 67:93-104

Mortensen, J., Bendtsen, J., Motyka, R.J., Lennert, K., Truffer, M., Fahnestock, M. and Rysgaard, S. 2013. On the seasonal freshwater stratification in the proximity of fast-flowing tidewater outlet glaciers in a sub-Arctic sill fjord. Journal of Geophysical Research Oceans 118:1-14. Doi:10.1002/jgrc.20134.

Parmentier, F.J.W., Christensen, T.R., Sørensen, L.L., Rysgaard, S., McGuire, A.D., Miller, P.A. and Walker, D.A. 2013. The impact of a lower sea-ice extent on Arctic greenhouse-gas exchange. Nature Climate Change 3:195-202. Doi:10.1038/nclimate1784.

Søgaard, D.H., Thomas, D., Rysgaard, S., Glud, R.N., Norman, L., Kaartokallio, H., Juul-Pedersen, T. and Geilfus, N-X. 2013. The relative contributions of biological and abiotic processes to the carbon dynamics in subarctic sea ice during the transition from winter to spring. Polar Biology. DOI 10.1007/s00300-013-1396-3.

Westergaard-Nielsen, A., Lund, M., Hansen, B.U. and Tamstorf, M.P. 2013. Camera derived vegetation greenness index as proxy for gross primary production in a low Arctic wetland area. ISPRS Journal of Photogrammetry and Remote Sensing 86:89-99.

Zamora-Terol, S., Nielsen, T.G. and Saiz, E. 2013. Plankton community structure and role of Oithona similis on the western coast of Greenland during the winter-spring transition. Marine Ecology Progress Series 483:85-102. 


Arendt, K.E., Juul-Pedersen, T., Mortensen, J., Blicher, M.E. and Rysgaard, S. 2012. A 5-year study of seasonal patterns in mesozooplankton community structure in a sub-Arctic fjord reveals dominance of Microsetella norvegica (Crustacea, Copepoda). Journal of Plankton Research 35(1):105-120.

Bluhm, B.A., Grebmeier, J.M., Archambault, P., Blicher, M.E., Guðmundsson, G., Iken, K., Lindal Jørgensen, L. and Mokievsky, V. 2012. Benthos In Arctic Report Card 2012 – tracking recent environmental changes (

Carlsson, P., Cornelissen, G., Bøggild, C.E., Rysgaard, S., Mortensen, J. and Kallenborn, R. 2012. Hydrology-linked spatial distribution of pesticides in a fjordsystem in Greenland. Journal of Environmental Monitoring 14(5):1437-1443.

Hansen, J.H., Hedeholm, R.B., Sunksen, K., Christensen, J.T. and Grønkjær, P. 2012. Spatial variability of carbon (delta C-13) and nitrogen (delta N-15) stable isotope ratios in an Arctic marine food web. Marine Ecology Progress Series 467:47-59.

Hedeholm, R., Grønkjær, P. and Rysgaard, S. 2012. Feeding ecology of capelin (Mallotus villosus Müller) in West Greenlandic waters. Polar Biology. Doi:10.1007/s00300-012-1193-4.

Heide-Jørgensen, M.P., Laidre, K.L., Hansen, R.G., Burt, M.L., Simon, M., Borchers, D.L., Hansen, J., Harding,K., Rasmussen, M., Dietz, R. and Teilmann, J. 2012. Rate of increase and current abundance of humpback whales in West Greenland. Journal of Cetacean Research and Management 12(1):1-14.

Krause-Jensen, D., Marbà, N., Olesen, B., Sejr, M.K., Christensen, P.B., Rodrigues, J., Renaud, P.E., Balsby, T.J. and Rysgaard, S. 2012. Seasonal sea ice cover as principal driver of spatial and temporal variation in depth extension and annual production of kelp in Greenland. Global Change Biology 18(10):2981-2994.

Long, M.H., Koopmans, D., Berg, P., Rysgaard, S., Glud, R.N. and Søgaard, D.H. 2012. Oxygen exchange and ice melt measured at the ice-water interface by eddy correlation. Biogeosciences 9:1-11. Doi:10.5194/bg-9-1-2012.

MacBride, J.R. and Douhovnikoff, V. 2012. Characteristics of the urban forests in arctic and near-arctic cities. Urban Forestry & Urban Greening 11(2):113-119.

Middelboe, M., Glud, R.N. and Sejr, M.K. 2012. Bacterial carbon cycling in a subarctic fjord: A seasonal study on microbial activity, growth efficiency, and virus-induced mortality in Kobbefjord, Greenland. Limnology Oceanography 57(6):1732-1742.

Rysgaard, S., Glud, R.N., Lennert, K., Cooper, M., Halden, N., Leaky, R., Hawthorne, F.C. and Barber, D. 2012. Ikaite crystals in melting sea ice leads to low pCO2 levels and high pH in Arctic surface waters. The Cryosphere 6:1-8. Doi: 10.5194/tc-6-901-2012.

Rysgaard, S., Mortensen, J., Juul-Pedersen, T., Sørensen, L.L., Lennert, K., Søgaard, D.H., Arendt, K.E., Blicher, M.E., Sejr, M.K. and Bendtsen, J. 2012. High air-sea CO2 uptake rates in nearshore and shelf areas of Southern Greenland: Temporal and spatial variability. Marine Chemistry 128:26-33.

Sealy, S.G. and Carter, H. R. 2012. Rare Inter-ocean Vagrancy in Crested Auklet and Parakeet Auklet. Waterbirds 35(1):64-73.

Simon, M., Johnson, M. and Madsen, P.T. 2012. Keeping momentum with a mouthful of water: behavior and kinematics of humpback whale lunge feeding. The Journal of Experimental Biology 215:3786-3798. Doi:10.1242/jeb.071092.

Søndergaard, J., Rigét, F., Tamstorf, M.P. and Larsen, M.M. 2012. Mercury transport in low-Arctic river in Kobbefjord, West Greenland. Water, Air and Soil Pollution 223(7):4333-4342.

Versteegh, E.A.A, Blicher, M.E., Mortensen, J., Rysgaard, S., Als, T.D. and Wanamaker Jr, A.D. 2012. Oxygen isotope ratios in the shell of Mytilus edulis: Archives of glacier melt water in Greenland?Biogeochemical Discussions 9, 12019-12046.


Agersted, M.D., Nielsen, T.G., Munk, P., Vismann, B. and Arendt, K.E. 2011. The functional biology and trophic role of krill (Thysanoessa raschii) in a Greenlandic fjord. Marine Biology 158(6):1387–1402. DOI 10.1007/s00227-011-1657-z.

Arendt, K.E., Dutz, J., Jonasdottir, S.H., Jung-Madsen, S., Mortensen, J., Moeller, E.F. and Nielsen, T.G. 2011. Effects of suspended sediments on copepods feeding in a glacial influenced sub-Arctic fjord. Journal of Plankton Research. DOI:10.1093/plankt/fbr054.

Blicher, M.E., Rasmussen, L.M., Sejr, M.K., Merkel, F.R. and Rysgaard, S. 2011. Abundance and energy requirements of eiders (Somateria spp.) suggest high predation pressure on macrobenthic fauna in a key wintering habitat in SW Greenland. Polar Biology 34(8):1105-1116.

Calbet, A., Riisgaard, K., Saiz, E., Zamara, S., Stedmon, C. and Nielsen, T.G. 2011. Phytoplankton growth and microzooplankton grazing along a sub-Arctic fjord (Godthåbsfjord, west Greenland). Marine Ecology Progress Series 434:77-99.

Hansen, E.S. 2011. A contribution to the lichen flora of the Kobbefjord area, West Greenland. Botanica Lithuanica 17(4):151-156.

Hedeholm, R., Grønkjær, P. and Rysgaard, S. 2011. Energy content and fecundity of capelin (Mallotus villosus) along a 1500-km latitudinal gradient. Marine Biology. DOI 10.1007/s00227-011-1651-5.

Mortensen, J., Lennert, K., Bendtsen, J. and Rysgaard, S. 2011. Heat sources for glacial melt in a sub-Arctic fjord (Godthåbsfjord) in contact with the Greenland Ice Sheet. Journal of Geophysical Research 116 - Oceans:116. Article Number: C01013. DOI: 10.1029/2010JC006528

Richter, A., Rysgaard, S., Dietrich, R., Mortensen, J. and Petersen, D. 2011. Coastal tides in West Greenland derived from tide gauge records. Ocean Dynamics 61:39-49.

Rysgaard, S., Bendtsen, J., Delille, B., Dieckmann, G., Glud, R.N., Kennedy, H., Mortensen, J., Papadimitriou, S., Thomas, D. and Tison, J-L. 2011. Sea ice contribution to air-sea CO2 exchange in the Arctic and Southern Oceans. Tellus 63B, 823-830. Doi: 10.1111/j.1600-0889.2011.00571.x.

Søgaard, D.H., Hansen, P.J., Rysgaard, S. and Glud, R.N. 2011. Growth limitation of three Arctic sea ice algal species: effects of salinity, pH, and inorganic carbon availability. Polar Biology. DOI 10.1007/s00300-011-0976-3.

Tang, K.W., Glud, R.N., Glud, A., Rysgaard, S. and Nielsen, T.G. 2011. Copepod guts as biogeochemical hotspots in the sea: evidence from microelectrode profiling of Calanus spp. Limnology and Oceanography 56:666-672.

Tang, K.W., Nielsen, T.G., Munk, P., Mortensen, J., Møller, E.F., Arendt, K.E., Tönnesson, K. and Juul-Pedersen, T. 2011. Metazooplankton community structure, feeding rate estimates, and hydrography in a meltwater-influenced Greenlandic fjord. Marine Ecology Progress Series 434:77-90.


Arendt, K.E., Nielsen, T.G., Rysgaard, S. and Tönneson, K. 2010. Differences in plankton community structure along Godthåbsfjord; from the Greenland Ice Sheet to offshore waters, SW Greenland. Marine Ecology Progress Series 401:49-62.

Blicher, M.E., Clemmesen, C., Rysgaard, S., Sejr, M.K. and Mempel, H. 2010. Seasonal and spatial variations in the RNA:DNA ratio and its relation to growth for sub-Arctic scallops. Marine Ecology Progress Series 407:87-98.

Blicher, M.E., Sejr, M.K. and Rysgaard, S. 2010. Seasonal growth variation of Chlamys islandica (Bivalvia) in the sub-Arctic Greenland is linked to food availability and temperature. Marine Ecology Progress Series 407:71-86.

Boye, T.K., Simon, M. and Madsen, P.T. 2010. Habitat use of humpback whales in Godthåbsfjord, West Greenland, with implications for commercial exploitation. Journal of the Marine Biological Association of the United Kingdom. doi:10.1017/S0025315410000755.

Glud, R.N., Berg, P., Hume, A., Batty, P., Blicher, M.E., Lennert, K. and Rysgaard, S. 2010. Benthic O2 exchange rates across hard-bottom sediments as quantified by eddy correlation in a sub-Arctic fjord system. Marine Ecology Progress Series 417:1-12.

Hedeholm, R., Grønkjær, P., Rosing-Asvid, A. and Rysgaard, S. 2010. Variation in size and growth of West Greenland capelin (Mallotus villosus) along a latitudinal gradient. ICES Journal of Marine Science. Doi:10.1093/icesjms/fsq024.

Mikkelsen, D.M. and Witkowski, A. 2010. Melting sea ice for taxonomic analysis: a comparison of four melting procedures. Polar Research 29:451-454.

Piña-Ochoa, E., Høgslund, S., Geslin, E., Cedhagen, T., Revsbech, N.P., Nielsen, L.P., Schweiger, M., Jorissen, F., Rysgaard, S. and Risgaard-Petersen, N. 2010. Widespread occurrence of nitrate storage and denitrification among Foraminifera and Gromiida. Proceedings of the National Academy of Sciences 107:1148-1153.

Post, E., Forchhammer, M.C., Tyler, N.J.C., Høye, T.T., Tamstorf, M.P., Christensen, T.R., Elberling, B., Gilg, O., Hik, D.S., Ims, R.A., Rysgaard, S., Stirling, I., Welker, J., Wookey, P.A., Schmidt, N.M. and Aastrup, P. 2010. Response to B. Griffith et al.'s E-Letter. 13 January 2010. Science.

Ruiz-Halpern, S., Sejr, M.K., Duarte, C.M., Krause-Jensen, D., Dalsgaard, T., Dachs, J. and Rysgaard, S. 2010. Air–water exchange and vertical profiles of organic carbon in a subarctic fjord. Limnology and Oceanography 55:1733-1740.

Sejr, M.K., Wlodarska-Kowalczuk, M., Legezynska, J. and Blicher, M.E. 2010. Macrobenthic species composition and diversity in the Godthaabsfjord system, SW Greenland. Polar Biology 33(4):421-431.   

Søgaard, D.H., Kristensen, M., Rysgaard, S. and Glud, R.N. 2010. Dynamics of autotrophic and heterotrophic activity in Arctic first-year sea ice: Case study from Marlene Bight, SW Greenland. Marine Ecology Progress Series 419:31-45.


Blicher, M.E., Sejr, M.K. and Rysgaard, S. 2009. High carbon demand of dominant macrozoobenthic species indicates their central role in ecosystem carbon flow in a sub-arctic fjord. Marine Ecology Progress Series 383:127-140.

Glud, R.N., Wöfel, J., Karsten, U., Kühl, M. and Rysgaard, S. 2009. Benthic microalgal production in the Arctic: Applied methods and status of the current database. Botanica Marina 52:559-571.

Rysgaard, S., Bendtsen, J.B., Pedersen, L.T., Ramløv, H. and Glud, R.N. 2009. Increased CO2 growth and decay in the Nordic Seas. Journal of Geophysical Research Ocean 114. C09011. Doi: 10.1029/2008JC005088.

Sejr, M.K., Blicher, M.E. and Rysgaard, S. 2009. Sea ice cover affects inter-annual and geographic variation in growth of the Arctic cockle uptake due to sea-ice Clinocardium ciliatum (Bivalvia)in Greenland. Marine Ecology Progress Series 389:149-158.


Mikkelsen, D.M., Rysgaard, S. and Glud, R.N. 2008. Microalgal composition and primary production in Arctic sea ice: a seasonal study from Kobbefjord (Kangerluarsunnguaq), West Greenland. Marine Ecology Progress Series 368:65-74.

Rysgaard, S., Glud, R.N., Sejr, M.K., Blicher, M.E. and Stahl, H.J. 2008. Denitrification activity and oxygen dynamics in Arctic sea ice. Polar Biology 31:527-537.


Berg, P., Swaney, D., Rysgaard, S., Thamdrup, B. and Fossing, H. 2007. A fast numerical solution of the general mass-conservation equation for solids and solutes in aquatic sediments. Journal of Marine Research 65:317-343.

Blicher, M., Rysgaard, S. and Sejr, M. K. 2007. Growth and production of the sea urchin, Strongylocentrotus droebachiensis (O.F. Müller, 1776), in a high arctic fjord and growth along a climate gradient (64 – 77°N). Marine Ecology Progress Series 341:89-102.

Rysgaard, S., Glud, R. N., Sejr, M. K., Bendtsen, J. and Christensen, P. B. 2007. Inorganic carbon transport during sea ice growth and decay: A carbon pump in polar seas. Journal of Geophysical Research 112: C03016. doi: 10.1029/2006jc003572.

Rysgaard, S., Glud, R. N., Sejr, M. K., Blicher, M. E. and Stahl, H. J. 2007. Denitrification activity and oxygen dynamics in Arctic sea ice. Polar Biology 31:527-537.

Schmid, M.C., Risgaard-Petersen, N., van de Vossenberg, J., Langezaal, S., Kuypers, M.M.M., Lavik, G., Brandsma, J., Damsté, J.S., van der Zwaan, B., Hulth, S., Thamdrup, B., Canfield, D., Dalsgaard, T., Rysgaard, S., Sejr, M. K., Strous, M., op den Camp, H. J. M and Jetten, M. S. M. 2007. Anaerobic Ammonium Oxidizing Bacteria in Marine Environments: Widespread Occurrence but low Diversity. Environmental Microbiology 9(6):1476-1484.

Sejr, M. K., Nielsen, T.G., Rysgaard, S., Risgaard-Petersen, N., Sturluson, M. and Blicher, M. 2007. The fate of pelagic organic carbon and importance of the benthic-pelagic coupling in a shallow cove in Disko Bay, west Greenland. Marine Ecology Progress Series 341:75-88.


Cook, P., Wenzhöfer, F., Rysgaard, S., Galaktionov, O., Eyre, B., Cornwell, J., Huettel, M. and Glud, R. N. 2006. Quantifying denitrification in permeable sediments: Insights from a two-dimensional simulation analysis and experimental data. Limnology and Oceanography Methods 4:294-307.

Forchhammer, M. C., Rasch, M. and Rysgaard, S. 2006. A conceptual framework for monitoring climate effects and feedback in arctic ecosystems. NERO 1st Annual Report, 2008.

Roeselers, G., Norries, T. B., Castenholz, R. W., Rysgaard, S., Glud, R. N., Kühl, M. and Muyzer, G. 2006. Diversity of phototrophic bacteria in microbial mats from arctic hot springs (Greenland). Environmental Microbiology. Doi:10.1111/j.1462-2920.2006.01103.x.